Volume 4 describes 94 fossil shell species as follows: Class Polyplacophora is described with 20 species, divided over 7 genera and 4 families. Clas Monoplacophora is described with the genus Neopilina. Finally, the superorder Archaeogastropoda is described with 73 species, divided over 26 genera and 7 superfamilies.
Every species is accompanied by the original name and the book’s title where the author described it. This is followed by a large bibliography, the distinguishing characteristics, and, where possible, the original diagnosis. Also included is a list of the fossil beds and their temporal and spatial distribution. Finally. observations on similar species and comments on nomenclature and ecological spaces are given. There are alphabetic indices by species name and generic and subgeneric name. The bibliography lists 233 titles. Five new species are described: Chiton colligens, Neopilina pliocenica, Clanculus bonfittoi, Gibbula verae, and Skenea bogii.
Malacofauna Pliocenica Toscana isn't only a series dealing with Tuscany's Pliocene fossil shells: it starts from Tuscany's Pliocenic Malacofauna and links it to Miocene, Pliocene, Pleistocene European fossil shells and the current Malacofauna of the Mediterranean Sea.
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