Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera)

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Language: English

Contents

1. Introduction 1 

2. Adult and larval morphology of the Aeschnidiidae 3 
2.1 The larva 3 
2.2 The adult 6 

3. Systematic Palaeontology 15 

3.1 Taxa previously attributed to Aeschnidium 15 
Genus Aeschnidium Westwood, 1854 15 
Aeschnidium bubas Westwood, 1854 15 
Aeschnidium antiquum (Brodie, 1845) 23 
Genus Misofaeschnidium gen. n 25 
Misofaeschnidium densum (Hagen, 1862) 25 
Genus Sinaeschnidia Hong, 1965 stat. rest. 30 
Sinaeschnidia martinezdelclosi sp. n. 39 

3.2 Taxa previously attributed to Urogomphus 40 
Genus Urogomphus Handlirsch, 1906 40 
Urogomphus giganteus (Münster in Germar, 1839) 43 
Urogomphus (?) species undetermined . 44 
Genus Lithoaeschnidium Nel & Martínez-Delclòs, 1993 stat. rest. 45 
Lithoaeschnidium viohli Nel & Martínez-Delclòs, 1993 stat. rest. 45 
Genus Brunetaeschnidium gen. n 46 
Brunetaeschnidium nusplingensis (Bechly, 1998) comb. n 46 
Brunetaeschnidium sp. 46 

3.3 Other genera previously described 48 
Genus Aegyptidium Schlüter & Hartung, 1982. 48 
Aegyptidium aburasiensis Schlüter & Hartung, 1982 48 
Genus Aeschnidiella Zalesky, 1953. 51 
Aeschnidiella kabanovi Zalesky, 1953 51 
Genus Aeschnidiopsis Tillyard, 1917 51 
Aeschnidiopsis flindersiensis (Woodward, 1884) 51 
Genus Bergeriaeschnidia Nel, Bechly & Martínez-Delclòs, 1996. 53 
Bergeriaeschnidia inexpectata Nel et al., 1996 stat. rest. 53 
Genus Gigantoaeschnidium Nel & Martínez-Delclòs, 1993. 55 
Gigantoaeschnidium ibericus Nel & Martínez-Delclòs, 1993. 55 
Genus Iberoaeschnidium Nel & Martínez-Delclòs, 1993. 63 
Iberoaeschnidium conquensis Nel & Martínez-Delclòs, 1993. 63 
Genus Leptaeschnidium Pritykina, 1977 63 
Leptaeschnidium latum Pritykina, 1977 65 
Leptaeschnidium araripina (Carle & Wighton, 1990) comb. n 65 
Genus Lleidoaeschnidium Nel & Martínez-Delclòs, 1993. 72 
Lleidoaeschnidium valloryi Nel & Martínez-Delclòs, 1993 72 
Lleidoaeschnidium maculatum sp. n. 72 
Genus Malmaeschnidium Nel & Martínez-Delclòs, 1993. 74 
Malmaeschnidium mayeri Nel & Martínez-Delclòs, 1993. 75 
Genus Nannoaeschnidium Nel & Martínez-Delclòs, 1993. 75 
Nannoaeschnidium pumilio Nel & Martínez-Delclòs, 1993. 75 
Genus Santanoptera Martill & Nel, 1996 78 
Santanoptera gabbotti Martill & Nel, 1996 78 
Genus Solnhofenia Bechly, 2000. 78 
Solnhofenia stoebeneri Bechly, 2000 78 
Genus Tauropteryx Pritykina, 1993. 78 
Tauropteryx krassilovi Pritykina, 1993 80 
Genus Stylaeschnidium Zhang & Zhang, 2001. 80 
Stylaeschnidium rarum Zhang & Zhang, 2001. 80 
Genus Dracontaeschnidium Zhang & Zhang, 2001. 80 
Dracontaeschnidium orientale Zhang & Zhang, 2001 80 
Descriptions of new genera 80 
Genus Angloaeschnidium gen. n. 80 
Angloaeschnidium toyei sp. n 82 
Cf. Angloaeschnidium toyei 86 
Angloaeschnidium montreuili sp. n . 89 
? Angloaeschnidium lacaui sp. n. 92 
Genus Cooperaeschnidium gen. n. 94 
Cooperaeschnidium durandi sp. n. 95 
Genus Delclosaeschnidium gen. n. 102 
Delclosaeschnidium magnum sp. n. 103 
Genus Diastatopsaeschnidium gen. n. 107 
Diastatopsaeschnidium reneeheiko sp. n. 107 
Genus Jarzembowskiaeschnidium gen. n. 108 
Jarzembowskiaeschnidium polandi sp. n. 110 
Genus Kessleraeschnidium gen. n. 114 
Kessleraeschnidium simonae sp. n 114 
Genus Rossaeschnidium gen. n. 118 
Rossaeschnidium patriciae sp. n. 118 
Genus Coramaeschnidium gen. n 120 
Coramaeschnidium minimum sp. n. 120 
Genus Kimmeridgebrachypteraeschnidium gen. n. 122 
Kimmeridgebrachypteraeschnidium etchesi sp. n. 122 

3.5 Undetermined new genera and species 124 

3.6 List of genera and species of Aeschnidiidae 134 

4. Phylogenetic analyses 137 

4.1 Method 137 

4.2 Position of the Aeschnidiidae relative to the Anisoptera 137 
4.2.1 Previous works 137 
4.2.2 New analysis 139 
4.2.3 List of characters 139 
4.2.4 List of taxa 149 
4.2.4.1 Outgroups 149 
4.2.4.2 Ingroup 149 
4.2.5 Results 149 

4.3 Phylogenetic analysis of the Aeschnidiidae 151 
4.3.1 Previous works 151 
4.3.2 Excluded taxa 151 
4.3.3 Included taxa (in alphabetic order) 151 
4.3.4 Outgroups 152 
4.3.5 List of character states. 152 
4.3.6 Results 165 

5. Conclusions 169 

Acknowledgements 170 
References 171

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