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Treats 208 taxa comprising 155 species and 55 subspecies. Colour plates illustrate 975 specimens, depicting all species and subspecies and genitalia illustrations of male and female for all species. Genera covered include Hadena, Carpadia, Polia, Hadula, Saragosa, Laconobia, Hecatera, Lasionycta, Sideridis, Conisania, Tholera, Mythimna and Leucania. Volume 4 also includes four species and two subspecies new to science, ten species new to Europe, two new genera and many new synonyms and combinations.