ZooKeys 599: Taxonomic Revision of the Rock-Dwelling Door Snail Genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae)

About this book

Montenegrina Boettger, 1877 (Mollusca: Gastropoda: Clausiliidae) is one of the few obligate limestone-dwelling door snail genera that have territorially well-defined distribution areas in southeastern Europe. Its allopatric populations inhabit patchy karst outcrops which, in combination with limited active dispersal, provides favourable conditions for radiation into multiple, geographically separated, and morphologically diverse forms.

This paper provides a revision of all known Montenegrina taxa, as well as descriptions of 4 new species and 17 new subspecies. A junior homonym has been replaced by a new name. Montenegrina thus comprises 29 species and 106 subspecies, making it the second most speciose door snail genus in Europe. Of each taxon types or specimens from the type localities are figured, distribution maps and georeferenced locality data are provided.

Contents

Introduction   5
Methods   7
Systematic part   12
      Genus Montenegrina Boettger, 1877   12
            Montenegrina apfelbecki (Sturany, 1907)   15
            Montenegrina attemsi (Wagner, 1914)   17
                  Montenegrina attemsi attemsi (Wagner, 1914)   18
                  Montenegrina attemsi jakupicensis Fauer, 1993   18
            Montenegrina cattaroensis (Rossmässler, 1835)   19
                  Montenegrina cattaroensis cattaroensis (Rossmässler, 1835)   20
                  Montenegrina cattaroensis antivaricostata nom. n.   21
                  Montenegrina cattaroensis umbilicata (Boettger, 1879)   22
            Montenegrina chiasma Nordsieck, 1972   23
            Montenegrina dofleini (Wagner, 1928)   24
                  Montenegrina dofleini dofleini (Wagner, 1928)   24
                  Montenegrina dofleini fagorum Nordsieck, 1974   26
                  Montenegrina dofleini kastoriae Nordsieck, 1972   27
                  Montenegrina dofleini pinteri Nordsieck, 1974   28
                  Montenegrina dofleini prespaensis Nordsieck, 1988   29
                  Montenegrina dofleini sinosi Páll-Gergely, 2010   30
                  Montenegrina dofleini wagneri Szekeres, 2006   30
            Montenegrina drimmeri Fehér & Szekeres, 2006   31
            Montenegrina fuchsi Brandt, 1961   32
                  Montenegrina fuchsi fuchsi Brandt, 1961   32
                  Montenegrina fuchsi klemmi Brandt, 1962   34
                  Montenegrina fuchsi muranyii Fehér & Szekeres, 2006   35
                  Montenegrina fuchsi pallida Fauer, 1993   36
            Montenegrina grammica Nordsieck, 1988   36
                  Montenegrina grammica grammica Nordsieck, 1988   37
                  Montenegrina grammica erosszoltani ssp. n.   37
                  Montenegrina grammica improvisa ssp. n.   38
            Montenegrina haringae sp. n.   39
            Montenegrina helvola (Küster, 1860)   40
                  Montenegrina helvola helvola (Küster, 1860)   41
                  Montenegrina helvola carinata Eross & Szekeres, 1999   42
                  Montenegrina helvola magna Fehér & Szekeres, 2006   42
                  Montenegrina helvola ornata Eross & Szekeres, 1999   44
                  Montenegrina helvola pageti Brandt, 1962   44
            Montenegrina hiltrudae Nordsieck, 1972   45
                  Montenegrina hiltrudae hiltrudae Nordsieck, 1972   45
                  Montenegrina hiltrudae desaretica ssp. n.   47
                  Montenegrina hiltrudae costulata Eross & Szekeres, 2006   48
                  Montenegrina hiltrudae dennisi Gittenberger, 2002   48
                  Montenegrina hiltrudae densicostulata Nordsieck, 1974   49
                  Montenegrina hiltrudae fusca Fehér & Szekeres, 2006   51
                  Montenegrina hiltrudae maasseni Gittenberger, 2002   51
                  Montenegrina hiltrudae protruda Gittenberger, 2002   52
                  Montenegrina hiltrudae robusta Nordsieck, 2009   53
                  Montenegrina hiltrudae sattmanni Nordsieck, 1988   53
                  Montenegrina hiltrudae selcensis ssp. n.   54
            Montenegrina janinensis (Mousson, 1859)   55
            Montenegrina laxa (Küster, 1861)   56
                  Montenegrina laxa laxa (Küster, 1861)   57
                  Montenegrina laxa dedovi Nordsieck, 2009   61
                  Montenegrina laxa delii ssp. n.   61
                  Montenegrina laxa disjuncta Fehér & Szekeres, 2006   62
                  Montenegrina laxa errans Eross & Szekeres, 2006   62
                  Montenegrina laxa iba Nordsieck, 1972   63
                  Montenegrina laxa kontschani Eross & Szekeres, 2006   64
                  Montenegrina laxa lakmosensis Nordsieck, 2009   65
                  Montenegrina laxa miraka Nordsieck, 1996   65
            Montenegrina lillae sp. n.   66
            Montenegrina minuscula Eross & Szekeres, 2006   67
            Montenegrina nana Fehér & Szekeres, 2006   68
                  Montenegrina nana nana Fehér & Szekeres, 2006   68
                  Montenegrina nana barinai ssp. n.   69
                  Montenegrina nana gracilis Eross & Szekeres, 2006   70
            Montenegrina okolensis Szekeres, 2006   71
                  Montenegrina okolensis okolensis Szekeres, 2006   71
                  Montenegrina okolensis caesia Fehér & Szekeres, 2006   72
            Montenegrina perstriata (Wagner, 1919)   74
                  Montenegrina perstriata perstriata (Wagner, 1919)   74
                  Montenegrina perstriata callistoma Fehér & Szekeres, 2006   75
                  Montenegrina perstriata diminuta Fehér & Szekeres, 1999   77
                  Montenegrina perstriata drimica Nordsieck, 1972   78
                  Montenegrina perstriata mavrovoensis Nordsieck, 2009   79
                  Montenegrina perstriata occidentalis Nordsieck, 1977   80
                  Montenegrina perstriata ochridensis (Wagner, 1925)   80
                  Montenegrina perstriata plenostoma Fehér & Szekeres, 2006   81
                  Montenegrina perstriata radikae Nordsieck, 1972   82
                  Montenegrina perstriata steffeki Eross & Szekeres, 2006   83
                  Montenegrina perstriata subcristatula Nordsieck, 1977   83
                  Montenegrina perstriata tenebrosa Nordsieck, 2009   84
            Montenegrina prokletiana sp. n.   85
                  Montenegrina prokletiana prokletiana ssp. n.   85
                  Montenegrina prokletiana kovacsorum ssp. n.   86
            Montenegrina rugilabris (Mousson, 1859)   87
                  Montenegrina rugilabris rugilabris (Mousson, 1859)   88
                  Montenegrina rugilabris edmundi Szekeres, 2006   88
                  Montenegrina rugilabris golikutensis ssp. n.   89
                  Montenegrina rugilabris gregoi ssp. n.   91
                  Montenegrina rugilabris irmengardis Klemm, 1962   93
                  Montenegrina rugilabris lambdaformis Reischütz & Sattmann, 1990   93
                  Montenegrina rugilabris welterschultesi Fehér & Szekeres, 1999   94
            Montenegrina skipetarica (Soós, 1924)   95
                  Montenegrina skipetarica skipetarica (Soós, 1924)   95
                  Montenegrina skipetarica csikii Eross & Szekeres, 2006   97
                  Montenegrina skipetarica danyii ssp. n.   98
                  Montenegrina skipetarica ersekensis Nordsieck, 1996   99
                  Montenegrina skipetarica flava Eross & Szekeres, 2006   100
                  Montenegrina skipetarica gurelurensis ssp. n.   101
                  Montenegrina skipetarica konitsae Nordsieck, 1972   102
                  Montenegrina skipetarica nobilis Eross & Szekeres, 2006   102
                  Montenegrina skipetarica pifkoi ssp. n.   103
                  Montenegrina skipetarica pindica Nordsieck, 1988   104
                  Montenegrina skipetarica puskasi ssp. n.   105
                  Montenegrina skipetarica remota Fehér & Szekeres, 2006   106
                  Montenegrina skipetarica rugosa Fehér & Szekeres, 2006   106
                  Montenegrina skipetarica thysi Loosjes & Loosjes-van Bemmel, 1988   107
                  Montenegrina skipetarica voidomatis Nordsieck, 1974   108
            Montenegrina soosi Eross & Szekeres, 2006   108
            Montenegrina sporadica Nordsieck, 1974   109
                  Montenegrina sporadica sporadica Nordsieck, 1974   110
                  Montenegrina sporadica tropojana ssp. n.  111
            Montenegrina stankovici (Urbanski, 1960)   111
            Montenegrina sturanyana sp. n.   112
                  Montenegrina sturanyana sturanyana ssp. n.   113
                  Montenegrina sturanyana gropana ssp. n.   114
                  Montenegrina sturanyana ostrovicensis ssp. n.   114
            Montenegrina subcristata (Pfeiffer, 1848)   116
            Montenegrina timeae Eross & Szekeres, 2006   122
            Montenegrina tomorosi Brandt, 1961   123
                  Montenegrina tomorosi tomorosi Brandt, 1961   124
                  Montenegrina tomorosi ampla Fehér & Szekeres, 2006   125
                  Montenegrina tomorosi coerulescens Nordsieck, 1996   126
                  Montenegrina tomorosi hunyadii ssp. n.   127
            Montenegrina zilchi Nordsieck, 1974   127
Discussion   128
Acknowledgements   131
References   132

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